Ray Romano
Manny (voice)
Acting
John Leguizamo
Sid (voice)
Acting
Denis Leary
Diego (voice)
Acting
Queen Latifah
Ellie (voice)
Acting
Seann William Scott
Crash (voice)
Acting
Josh Peck
Eddie (voice)
Acting
Jay Leno
Fast Tony (voice)
Acting
Will Arnett
Lone Gunslinger Vulture (voice)
Acting
Chris Wedge
Scrat (voice)
Acting
Peter Ackerman
Dung Beetle Dad (voice)
Writing
Caitlin Rose Anderson
Glypto Boy Billy / Beaver Girl (voice)
Acting
Connor Anderson
Rhino Boy / Beaver Boy (voice)
Acting
Joseph Bologna
Mr. Start (voice)
Acting
Jack Crocicchia
Elk Boy (voice)
Acting
Peter DeSève
Condor Chick (voice)
Visual Effects
Ariel Winter
Additional Voices (voice)
Acting
Clea Lewis
Female Mini Sloth (voice)
Acting
Stephen Root
Aardvark Dad (voice)
Acting
Nicole DeFelice
Start Girl (voice)
Acting
Debi Derryberry
Diatryma Mom (voice)
Acting
Marshall Efron
Start Dad (voice)
Acting
Tom Fahn
Stu / Male Ox (voice)
Acting
Jason Fricchione
Molehog Grandpa / Shovelmouth Male (voice)
Acting
George Jacobs
Beaver Dad (voice)
Acting
Brian Scott McFadden
Glyptodon (voice)
Acting
Jansen Panettiere
Shovelmouth Boy (voice)
Acting
Gregory Romano
Start Boy 1 (voice)
Acting
Matt Romano
Start Boy 2 (voice)
Acting
Carlos Saldanha
Dodo (voice)
Directing
Manoela Scarpa Saldanha
Molehog Kid 1 (voice)
Acting
Sofia Scarpa Saldanha
Molehog Kid 2 (voice)
Acting
Reyna Shaskan
Diego's Bird Girl (voice)
Acting
James Sie
Freaky Male (voice)
Acting
Cindy Slattery
Aardvark Mom (voice)
Acting
Mindy Sterling
Female Ox (voice)
Acting
Renée Taylor
Mrs. Start (voice)
Acting
Alan Tudyk
Cholly (voice)
Acting
Claudia Besso
Additional Voices (voice)
Acting
Madeleine Martin
Additional Voices (voice)
Acting
Sherry Lynn
Additional Voices (voice)
Acting
Mickie McGowan
Additional Voices (voice)
Acting
Emily Anderson
Additional Voices (voice)
Acting
Jack Angel
Additional Voices (voice)
Acting
Shane Baumel
Additional Voices (voice)
Acting
Bob Bergen
Additional Voices (voice)
Acting
Kyle J. Bogert
Additional Voices (voice)
Acting
Rodger Bumpass
Additional Voices (voice)
Acting
Paul Butcher
Additional Voices (voice)
Acting
Blanca Camacho
Additional Voices (voice)
Acting
John Cygan
Additional Voices (voice)
Acting
Jennifer Darling
Additional Voices (voice)
Acting
Olivia DeLaurentis
Additional Voices (voice)
Acting
Harrison Fahn
Additional Voices (voice)
Acting
Bill Farmer
Additional Voices (voice)
Acting
Crispin Freeman
Additional Voices (voice)
Acting
Tim Kirkpatrick
Additional Voices (voice)
Acting
David H. Kramer
Additional Voices (voice)
Production
Danny Mann
Additional Voices (voice)
Acting
Laura Marano
Additional Voices (voice)
Acting
Matt McCarthy
Additional Voices (voice)
Acting
Laraine Newman
Additional Voices (voice)
Acting
Jan Rabson
Additional Voices (voice)
Acting
Zack Shada
Additional Voices (voice)
Acting
Thomas Sharkey
Additional Voices (voice)
Acting
Ross Simanteris
Additional Voices (voice)
Acting
Rachel Stand
Additional Voices (voice)
Acting
Spencer Lacey Ganus
Additional Voices (voice)
Acting
Jess Harnell
Additional Voices (voice)
Acting
Alex Sullivan
James (voice)
Acting
James Godwin
Traffic Vulture
Acting